俱舍論疏 3.136

3.136

論「別有男相至男身所有」者，此通經。雖無男根，而有男身眾相，故名男也。非如欲界無男根即無男相。

Regarding the commentary's statement: "There are distinct male characteristics [...] possessed by a male body": this resolves [the conflict with] the sūtra. Although they lack the male faculty, they possess the various characteristics of a male body; therefore they are called male. It is not as in the desire realm, where lacking the male faculty means lacking the male characteristics.