瑜伽論記 3.237

3.237

二十、苦及苦因，五能生苦者，謂無明行愛取有。苦胎藏五者，謂識等五種子故，苦體即是生及老死。

Regarding 'Twentieth, suffering and the causes of suffering: five can generate suffering,' these refer to ignorance, volitional formations, craving, clinging, and existence. 'The five that are the matrix of suffering' refer to the seeds of the five beginning with consciousness; the substance of suffering is precisely birth and aging-and-death.